This inherited property is initialized in the InitializeTreeVertexValues method. A value of Spot.Default will cause SetPortSpots to assign a FromSpot based on the parent node's TreeVertex.Angle. If the value is other than NoSpot, it is just assigned. When Path is TreePath.Source, the port's ToSpot is set instead of the FromSpot.
This sets the AlternateDefaults's property of the same name.